---
title: "Flying and gliding animals"
chunk: 4/8
source: "https://en.wikipedia.org/wiki/Flying_and_gliding_animals"
category: "reference"
tags: "science, encyclopedia"
date_saved: "2026-05-05T03:46:37.316378+00:00"
instance: "kb-cron"
---

Unlike most air vehicles, in which the objects that generate lift (wings) and thrust (engine or propeller) are separate and the wings remain fixed, flying animals use their wings to generate both lift and thrust by moving them relative to the body. This has made the flight of organisms considerably harder to understand than that of vehicles, as it involves varying speeds, angles, orientations, areas, and flow patterns over the wings.
A bird or bat flying through the air at a constant speed moves its wings up and down (usually with some fore-aft movement as well). Because the animal is in motion, there is some airflow relative to its body which, combined with the velocity of its wings, generates a faster airflow moving over the wing. This will generate lift force vector pointing forwards and upwards, and a drag force vector pointing rearwards and upwards. The upwards components of these counteract gravity, keeping the body in the air, while the forward component provides thrust to counteract both the drag from the wing and from the body as a whole. Pterosaur flight likely worked in a similar manner, though no living pterosaurs remain for study.
Insect flight is considerably different, due to their small size, rigid wings, and other anatomical differences. Turbulence and vortices play a much larger role in insect flight, making it even more complex and difficult to study than the flight of vertebrates. There are two basic aerodynamic models of insect flight. Most insects use a method that creates a spiralling leading edge vortex. Some very small insects use the fling-and-clap or Weis-Fogh mechanism in which the wings clap together above the insect's body and then fling apart. As they fling open, the air gets sucked in and creates a vortex over each wing. This bound vortex then moves across the wing and, in the clap, acts as the starting vortex for the other wing. Circulation and lift are increased, at the price of wear and tear on the wings.

== Limits and extremes ==

=== Flying and soaring ===

==== Largest ====
The largest known flying animal was formerly thought to be Pteranodon, a pterosaur with a wingspan of up to 7.5 metres (25 ft). However, the more recently discovered azhdarchid pterosaur Quetzalcoatlus is much larger, with estimates of the wingspan ranging from 9 to 12 metres (30 to 39 ft). Some other recently discovered azhdarchid pterosaur species, such as Hatzegopteryx, may have also wingspans of a similar size or even slightly larger. Although it is widely thought that Quetzalcoatlus reached the size limit of a flying animal, the same was once said of Pteranodon. The heaviest living flying animals are the kori bustard and the great bustard with males reaching 21 kilograms (46 lb). The wandering albatross has the greatest wingspan of any living flying animal at 3.63 metres (11.9 ft). Among living animals which fly over land, the Andean condor and the marabou stork have the largest wingspan at 3.2 metres (10 ft). Studies have shown that it is physically possible for flying animals to reach 18-metre (59 ft) wingspans, but there is no firm evidence that any flying animal, not even the azhdarchid pterosaurs, got that large.

==== Smallest ====
There is no minimum size for getting airborne. Indeed, there are many bacteria floating in the atmosphere that constitute part of the aeroplankton. However, to move about under one's own power and not be overly affected by the wind requires a certain amount of size. The smallest flying vertebrates are the bee hummingbird and the bumblebee bat, both of which may weigh less than 2 grams (0.071 oz). They are thought to represent the lower size limit for endotherm flight. The smallest flying invertebrate is a fairyfly wasp species, Kikiki huna, at 0.15 mm (0.0059 in) (150 μm).

==== Fastest ====
The fastest of all known flying animals is the peregrine falcon, which when diving travels at 300 kilometres per hour (190 mph) or faster. The fastest animal in flapping horizontal flight may be the Mexican free-tailed bat, said to attain about 160 kilometres per hour (99 mph) based on ground speed by an aircraft tracking device; that measurement does not separate any contribution from wind speed, so the observations could be caused by strong tailwinds.

==== Slowest ====
Most flying animals need to travel forward to stay aloft. However, some creatures can stay in the same spot, known as hovering, either by rapidly flapping the wings, as do hummingbirds, hoverflies, dragonflies, and some others, or carefully using thermals, as do some birds of prey. The slowest flying non-hovering bird recorded is the American woodcock, at 8 kilometres per hour (5.0 mph).

==== Highest flying ====
There are records of a Rüppell's vulture Gyps rueppelli, a large vulture, being sucked into a jet engine 11,550 metres (37,890 ft) above Côte d'Ivoire in West Africa. The animal that flies highest most regularly is the bar-headed goose Anser indicus, which migrates directly over the Himalayas between its nesting grounds in Tibet and its winter quarters in India. They are sometimes seen flying well above the peak of Mount Everest at 8,848 metres (29,029 ft).

=== Gliding and parachuting ===

==== Most efficient glider ====
This can be taken as the animal that moves most horizontal distance per metre fallen. Flying squirrels are known to glide up to 200 metres (660 ft), but have measured glide ratio of about 2. Flying fish have been observed to glide for hundreds of metres on the drafts on the edge of waves with only their initial leap from the water to provide height, but may be obtaining additional lift from wave motion. On the other hand, albatrosses have measured lift–drag ratios of 20, and thus fall just 1 meter for every 20 in still air.